ForBio and UiB Dead Wood Course 2021

[This course has been tentatively rescheduled to October 2021 due to the COVID-2019 pandemic. The course is 5 days long with dates to be announced later. Application will be re-opened in 2021 following the national Norwegian guidelines on international travel]


The course is organized by the University Museum of Bergen, the Lomonosov Moscow State University, the Norwegian University of Life Sciences, the Norwegian Institute for Nature Research, the University of Oslo, and the Research School in Biosystematics – ForBio, with the support from the University of Helsinki, and funding from the Norwegian Agency for International Cooperation and Quality Enhancement in Higher Education (Diku).


The intensive five-day course program is comprised by lectures, workshops, excursions, and an obligatory book exam. The target group is advanced master and PhD students. It is recommended that students have taken basic courses in entomology, mycology, and ecology. The aim of the course is to familiarize students with various aspects of the habitat, organism groups and ecological processes in dead wood. Dead wood experts will provide deeper coverage of their fields and research and will give examples of case studies and the workshop sessions. We will go through a broad range of topics, such as biodiversity in dead wood in boreal and temperate environments, biodiversity conservation and restoration, and of course all kinds of lignicolous organisms: fungi, insects, and more.

Assignment and credits

Course attendance is 2 ECTS. Student must choose one of the books for the obligatory course exam, which takes place on the last day of the course. Beginners and advanced beginners have to choose Afterlife of a Tree, 2 ECTS, or Biodiversity in Dead Wood, 3 ECTS. Students already familiar with dead wood systems may instead choose Ecology of Saprotrophic Basidiomycetes or Saproxylic Insects for an advanced 3 ECTS book exam.

  • Bobiec A, Gutowski JM, Zub K, Pawlaczyk P, Laudenslayer WF 2005. The Afterlife of a Tree. WWF Poland, 252 pp. ISBN 83-920712-1-2 (basic book, 2 credits)

  • Stokland JN, Siitonen J, & Jonsson BG 2012. Biodiversity in Dead Wood. Cambridge Univ. Press, 524 pp. ISBN 9780521717038 (intermediate level, 3 credits).

  • Lynne B, Frankland J, Van West P, eds. 2007. Ecology of saprotrophic basidiomycetes Elsevier, 386 pp. eBook ISBN 9780080551500 / ISBN 9780123741851 (advanced book on fungi, 3 credits).

  • Ulyshen MD, ed. 2018. Saproxylic Insects. Diversity, Ecology and Conservation. Springer, 904 pp. eBook ISBN 978-3-319-75937-1 / ISBN 978-3-319-75936-4 (advanced book on insects, 3 credits).


The course will take place at the Espegrend Marine Biological Station located in the proximity of the Bergen airport and within a short drive to the city centre. The station provides access to various managed and undisturbed forest biotopes and has excellent  teaching and accommodation facilities.  

Course leader

Dmitry Schigel - University of Helsinki


Will be announced in early 2021


Minimum 12, maximum 16 students; priority given to students from Norway and Russia, a few additional places for other countries. Participants will be selected based on the registration form, which includes a paragraph for motivation and importance of the course for your study/work (max 100 words).

Course fee

There is no course fee required from ForBio members or associates.  Find out about how to become a ForBio member/associate here.


Shared accommodation (double rooms with a private bathroom) - NOK150/night; bench fee - NOK50/day, bed linen and towels are provided - prices are from 2019 and are subject to change.

Financial support

ForBio members and associates from Norway and Russia are supported in full, including travel. 

Accommodation is covered for non-Norwegian ForBio members (i.e. MSc and PhD students from other Scandinavian countries).

Applicants outside Norway or Russia need to cover their travel and accommodation costs.

Meals are covered for all course participants.


Participants are requested to arrange own travel/medical insurance valid in Norway including cover of COVID-19 treatment.

Visa support

Accepted applicants will receive official invitation letters from the course organizers, visa and medical insurance costs are not covered.


English, with limited support in Russian.

Application will be open in 2021.


Dmitry Schigel ( - course programme and content
Nataliya Budaeva ( - ForBio coordinator - travel arrangements and visa questions


Published Dec. 9, 2019 9:23 PM - Last modified Aug. 20, 2020 10:32 AM